Black Sea biomarkers · 35 verified molecular structures
Multi-proxy molecular evidence for gradual limnic–marine transition at 8.5–8.0 ka · click any card to expand
Marine Archaea Ammonia-oxidizing Archaea Soil bacteria (Acidobacteriota) Haptophyte algae Terrestrial plants Universal / multiple
A Isoprenoidal GDGTs — Marine Group I.1a Thaumarchaeota 4 compounds · TEX₈₆ components · all 4 evidence tiers
B Crenarchaeol family — diagnostic of marine Thaumarchaeota 3 compounds · BIT denominator · Holzheimer 2021 total synthesis
C Branched GDGTs (5-methyl) — Acidobacteriota 9 compounds · MBT'5ME · BIT numerators · Mahapatra 2025 synthesis of Ia
D Branched GDGTs (6-methyl primed) 6 compounds · MBT'6ME · alkaline/dry-soil indicator
E Hydroxylated iGDGTs — ammonia-oxidizing archaea 3 compounds · RI-OH · axenic culture (Varma 2024)
F Non-GDGT biomarkers — alkenones, n-alkanes, archaeol, iso-diabolic 10 compounds · UK'₃₇ · ACL · CPI · diagnostic for haptophytes & higher plants